Research in writing

Dr. Fiorillo has published over 100 popular scientific papers.

Dr. Fiorillo's recent papers

1       Blob, R.W., and Fiorillo, A.R. 1996. The significance of microfossil size and shape distributions in faunal abundance reconstructions: a Late Cretaceous example. Paleobiology 22(3):422-435.

2       Fiorillo, A.R., and May, C.L. 1996.  Preliminary report on the taphonomy and depositional setting of a new dinosaur locality in the Morrison Formation (Brushy Basin Member) of Curecanti National Recreation Area, Colorado. in, The Continental Jurassic, Morales, M. (ed.). Mus. Northern Ariz. Bull. 60:555-561.

3       Fiorillo, A.R., and McCarty, D.K., 1996.  Paleopedological evidence for a humid paleoenvironment in the lower part of the Brushy Basin Member, Morrison Formation, of Curecanti National Recreation Area, Colorado. in, The Continental Jurassic, Morales, M. (ed.). Mus. Northern Ariz. Bull. 60:575-580.

4       Fiorillo, A.R. 1997.  Stratigraphic distribution of fossil vertebrates in the Judith River Formation (Upper Cretaceous) of Wheatland and Golden Valley Counties, south-central Montana. Northwest Geol. 27:1-12.

5       Fiorillo, A.R., and Weishampel, D.B.  1997.  Teeth, tooth wear and replacement. in, The Dinosaur Encyclopedia, Currie, P.J. and Padian, K. (eds.). Academic Press, San Diego, p. 743-745.

6       Fiorillo, A.R. 1997.  Taphonomy. in, The Dinosaur Encyclopedia, Currie, P.J. and Padian, K. (eds.). Academic Press, San Diego, p. 713-716.

7       Fiorillo, A.R. 1998. Bone modification features on sauropod remains (Dinosauria) from the Freezeout Hills Quarry N (Morrison Formation) of southeastern Wyoming and their contribution to fine-scale paleoenvironmental interpretation.  Modern Geology, 23:111-126.

8       Fiorillo, A. R.  1998. Notes on the Half Moon Canyon Section of the Beartooth Butte Formation (Lower Devonian), Big Snowy Mountains, central Montana.  Northwest Geology, 28:15-19.

33    Fiorillo, A.R. 1998. Preliminary report on a new sauropod locality in the Javelina Formation (Late Cretaceous), Big Bend National Park, Texas. in, Research in the National Parks, Santucci, V. (ed.), Geologic Resources Division Technical Report, NPS/NRGRD/GRDTR-98/01, p. 29-31.

34    Fiorillo, A.R. 1998.  Dental microwear patterns from the sauropod dinosaurs Camarasaurus and Diplodocus:  evidence for resource partitioning in the Late Jurassic of North America.  Historical Biology, 13:1-16.

35    Fiorillo, A.R. 1998. Measuring fossil reworking within a fluvial system: an example from the Hell Creek Formation of eastern Montana.  In, Advances in Vertebrate Paleontology and Geochronology, Tomida, Y., Flynn, L.J., and Jacobs, L.L. (eds.), National Science Museum Monographs, No. 14, Tokyo, Japan, p. 243-251.

36     Chure, D.J., Fiorillo, A.R., and Jacobsen, A. 1998.  Prey bone utilization by predatory dinosaurs in the Late Jurassic of North America with comments on prey bone use by dinosaurs throughout the Mesozoic. GAIA 15:227-232.

37    Fiorillo, A.R.  1999.  Determining the Relative Roles of Climate and Tectonics in the Formation of the Vertebrate Fossil Record: a Perspective from the Late Cretaceous of Western North America.  Records of the Western Australian Museum Suppl. 57:219-228.

38    Fiorillo, A.R. 1999.  Nonmammalian microvertebrate remains from the Robison Eggshell Site, Cedar Mountain Formation (Early Cretaceous), Emery County, Utah. In, Vertebrate Paleontology in Utah, Gillette, David D. (ed.), Utah Geological Survey Miscellaneous Pub. 99-1, p. 259-268.

39    Hasiotis, S.T., Fiorillo, A.R., and Hanna, R.R. 1999.  Borings in Jurassic dinosaur bones:  trace fossil evidence of beetle interactions with vertebrates. In, Vertebrate Paleontology in Utah, Gillette, David D. (ed.), Utah Geological Survey Miscellaneous Pub. 99-1, p. 193-200.

40    Fiorillo, A.R.  1999. Non-marine Trace Fossils from the Morrison Formation (Jurassic) of Curecanti National Recreation Area, Colorado.  In, V. Santucci, and L. McClelland (eds.), National Park Service Paleontological Research Volume, Technical Report NPS/NRGRD/GRDTR-99/03, p. 42-46.

41    Englemann, G., and Fiorillo, A.R.  2000. The taphonomy and paleoecology of the Morrison Formation determined from a field survey of fossil localities. GeoResearch Forum, 6:533-540.

42    Fiorillo, A.R., Padian, K., and Musikasinthorn, C. 2000.  Taphonomy and depositional setting of the Placerias Quarry (Chinle Formation:  Late Triassic, Arizona). PALAIOS 15:373-386.

43    Fiorillo, A.R.  2000.  The ancient environment of the Beartooth Butte Formation (Devonian) in Wyoming and Montana: combining paleontological inquiry with federal management needs.  in: Cole, David N. and McCool, Stephen F. eds. Proceedings: Wilderness Science in a Time of Change. Proc. RMRS-P-000. Ogden, UT: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station. v. 3, p. 160-167.

44    Fiorillo, A.R., and Gangloff, R.A.  2000.  Theropod teeth from the Prince Creek Formation (Cretaceous) of northern Alaska, with speculations on arctic dinosaur paleoecology.  Journal of Vertebrate Paleontology 20:675-682.

45    Fiorillo, A.R., Santucci, V.L., Gangloff, R.A., Armato, P.J., and Kucinski, R.  2001.  Establishing paleontological baseline data for research and management needs: lessons learned from the NPS Alaska Region. in: Santucci, V.L. and McClelland, L. eds. Proceedings of the 6th Conference on Fossil Resources on Public Lands. Geological Resources Division Technical Report NPS/NRGRD/GRDTR-01/01, p. 123-129.

46    Fiorillo, A.R., and Gangloff, R.A.  2001.  The caribou migration model for Arctic hadrosaurs (Ornithischia: Dinosauria): a reassessment.  Historical Biology 15:323-334.

47    Fiorillo, A.R. 2003.  Dinosaurs of Arctic Alaska.  The Polar Times v. 3, no. 3, p. 4-5.

48    Engelmann, G.F., Chure, D.J., and Fiorillo, A.R.  2004.  The implications of a dry climate for the paleoecology of the fauna of the Late Jurassic Morrison Formation.  Sedimentary Geology 167:297-308.

49    Fiorillo, A.R., and Eberth, D.A.  2004.  Dinosaur taphonomy.  In, Weishampel, D.B., Dodson, P., and Osmolska, H., eds., The Dinosauria, 2nd ed.  University of California Press, Berkeley.  Pp. 607-613.

50    Fiorillo, A.R., and Parrish, J.T.  2004. The first record of a Cretaceous dinosaur from western Alaska. Cretaceous Research 25:453-458.

51    Gangloff, R.A., Fiorillo, A.R., and D.W. Norton.  2005.  The first Pachycephalosaurine (Dinosauria) from the Paleo-Arctic and its paleogeographic implications.  Journal of Paleontology 79:997-1001.

52    Fiorillo, A.R.  2005.  Turtle tracks in the Judith River Formation (Upper Cretaceous) of south-central Montana.  Palaeontologica Electronica. v. 8, no. 1, 11p., 1 MB.

53    Fiorillo, A.R.  2006.  Review of the Dinosaur Record of Alaska with comments regarding Korean Dinosaurs as comparable high-latitude fossil faunas.  Journal of Paleontological Society of Korea, 22(1):15-27.

54    Fiorillo, A.R.  2006.  Microwear patterns of the teeth of Cretaceous theropod dinosaurs from Montana and Alaska with inferences about diet and ecology.  In, Barrett, P.M., and Evans, S.E. (eds.) 9th Symposium on Mesozoic Terrestrial Ecosystems.  Manchester, UK., Cambridge University Press, pp., 44-47.

55    Fiorillo, A.R.  2007.  Dinosaurs of Alaska: an update.  Polar Times, v. 3, no. 10, p. 32-33.

56    Eberth, D., Rogers, R.R., and Fiorillo, A.R.  2007.  A practical approach to the study of bonebeds.  In, Bonebeds: genesis, analysis, and paleobiological significance, R.R. Rogers, D. Eberth, A.R. Fiorillo (eds.).  University of Chicago Press, Chicago.  Pp. 265-331.

57    Jacobs, L., Fiorillo, A.R., Gangloff, R.A., and Pasch, A.  2007.  Desmostylian remains from Unalaska Island, Aleutian Chain, Alaska. In, Bulletin of the Carnegie Museum of Natural History, No. 39, Beard, K.C. and Z.-X. Luo (eds.), Carnegie Museum of Natural History, Pittsburgh.  Pp. 189-202.

58    Fiorillo, A.R., McCarthy, P.J. and Breithaupt, B., Brease, P.  2007.  Dinosauria and fossil Aves footprints from the Lower Cantwell Formation (latest Cretaceous), Denali Park and Preserve, Alaska.  Alaska Park Science, 6:41-43.

59    Fiorillo, A.R.  2008.  Lack of variability in feeding patterns of the sauropod dinosaurs Diplodocus and Camarasaurus (Late Jurassic, Western USA) with respect to climate as indicated by tooth wear features.  In, Vertebrate microfossil assemblages: their role in paleoecology and paleobiogeography.  Sankey, J.T. and Baszio, S. (eds.), University of Indiana Press, Bloomington. p. 104-116.

60    Fiorillo, A.R. 2008. On the occurrence of exceptionally large teeth of Troodon (Dinosauria:Saurischia) from the Late Cretaceous of northern Alaska.  PALAIOS, 23:322-328.

61    Fiorillo, A.R.  2008. Cretaceous dinosaurs of Alaska:  Implications for the origins of Beringia.  In, Blodgett, R.B., and Stanley, G. (eds.), The Terrane Puzzle: new perspectives on paleontology and stratigraphy from the North American Cordillera.  Geological Society of America Special Paper 442, p. 313-326, Boulder.

62    Myers, T.S., and Fiorillo, A.R. 2009.  Evidence for gregarious behaviour and age segregation in sauropod dinosaurs.  Palaeogeography, Palaeoclimatology, Palaeoecology, 274:196-204.

63    Fiorillo, A.R., Tykoski, R.S., Currie, P.J., McCarthy, P.J., and Flaig, P.  2009.  Description of Two Partial Troodon Braincases from the Prince Creek Formation (Upper Cretaceous), North Slope Alaska. Journal of Vertebrate Paleontology, 29:178-187.

64    Lindgren, J., Alwmark, C., Caldwell, M.W., and Fiorillo, A.R.  2009.  Skin of the Cretaceous mosasaur Plotosaurus: implications for aquatic adaptations in giant marine reptiles.  Biology Letters. 5(4):528-531.

65    Fiorillo, A.R., Hasiotis, S.T., Kobayashi, Y., and Tomsich, C.S. 2009.  A pterosaur manus track from Denali National Park, Alaska Range, Alaska, USA.  PALAIOS, 24:466-472.

66    Fiorillo, A.R., LePain, D.L., Decker, P.L., and McCarthy, P.J. 2009.  A probable ceratopsian manus track, Nanushuk Formation, Albian, northern Alaska.  In, Buscalioni, A.D., and Martinez, M.F., eds., 10th International Mesozoic Terrestrial Ecosystems and Biota, Teruel, Spain, Universidad Autonoma de Madrid, p. 87-88.

67    Chinsamy, A., Fiorillo, A.R., and Tumarkin-Deratzian, A.R.  2009.  Hadrosaurs from the North Slope of Alaska: biological insights from bone microstructure.  In, Buscalioni, A.D., and Martinez, M.F., eds., 10th International Mesozoic Terrestrial Ecosystems and Biota, Teruel, Spain, Universidad Autonoma de Madrid, p. 77-78.

68    Jacobs, L.J., Fiorillo, A.R., Nishida, Y., Fitzgerald, E.M.G.  2009.  Mid-Cenozoic mammals from Alaska.  In, Papers on geology, vertebrate paleontology, and biostratigraphy in honor of Michael O. Woodburne.  Albright, L.B. III (ed.) Museum of Northern Arizona Bulletin 65, Flagstaff.  Pp. 171-184.

69    Tykoski, R.S., and Fiorillo, A.R.  2010. An enantiornithine bird from the lower Middle Cenomanian of Texas.  Journal of Vertebrate Paleontology, 30:288-292.

70    Fiorillo, A.R., McCarthy, P.J., Flaig, P.P., Brandlen, E., Norton, D.W., Zippi, P., Jacobs, L., and Gangloff, R.A.  2010.  Paleontology and paleoenvironmental interpretation of the Kikak-Tegoseak Quarry (Prince Creek Formation: Late Cretaceous), northern Alaska: a multi-disciplinary study of a high-latitude ceratopsian dinosaur bonebed. In, New Perspectives on Horned Dinosaurs.  Ryan, M.J, Chinnery-Allgeier, B.J., and Eberth, D.A., (eds.), Indiana University Press, Bloomington.  Pp. 456-477.

71    Gangloff, R.A., and Fiorillo, A.R.  2010.  Taphonomy and paleoecology of a bonebed from the Prince Creek Formation, North Slope, Alaska.  PALAIOS, 25(5):299-317.

72    Fiorillo, A.R. and McCarthy, P.J.  2010.  Ancient polar ecosystems and environments: proxies for understanding climate change and global warming – an introduction.  Palaeogeography, Palaeoclimatology, Palaeoecology, 295:345-347.

73    Parrish, J.T., Fiorillo, A.R., Jacobs, B.F., Currano, E.D., and Wheeler, E.A.  2010.  The Ketavik Formation: A new stratigraphic unit and its implications for the Paleogene paleogeography and paleoclimate of southwestern Alaska.  Palaeogeography, Palaeoclimatology, Palaeoecology, 295:348-362.

74    Fiorillo, A.R., McCarthy, P.J., and Flaig, P.P.  2010.  Taphonomic and Sedimentologic Interpretations of the Dinosaur-Bearing Upper Cretaceous Strata of the Prince Creek Formation, Northern Alaska:  Insights from an Ancient High-Latitude Terrestrial Ecosystem.  Palaeogeography, Palaeoclimatology, Palaeoecology, 295:376-388.

75    Fiorillo, A.R., Decker, P.L., LePain, D.L., Wartes, M., and McCarthy, P.J.  2010.  A probable Neoceratopsian Manus Track from the Nanushuk Formation (Albian, Northern Alaska).  Journal of Iberian Geology 36(2):165-174.

76    Fiorillo, A.R., Hasiotis, S.T., Kobayashi, Y., Breithaupt, B.H., McCarthy, P.J.  2011.  Bird tracks for the Upper Cretaceous Cantwell Formation of Denali National Park, Alaska, USA: a new perspective on ancient polar vertebrate biodiversity.  Journal of Systematic Palaeontology 9(1):33-49.

77    Flaig, P., McCarthy, P.J., and Fiorillo, A.R.  2011.  A tidally-influenced, high-latitude alluvial/coastal plain: the Late Cretaceous (Maastrichtian) Prince Creek Formation, North Slope, Alaska.  In, From River to Rock Record: The Preservation of Fluvial Sediments and their Subsequent Interpretation.  North, C., Davidson, S., and Leleu, S. (eds.), SEPM Special Publication 97, p. 233-264.

78    Adams, T., and Fiorillo, A.R.  2011.  Platypterygius Huene, 1922 (Ichthyosauria: Ophthamosauridae) from the Upper Cretaceous of Texas.  Palaeontologia Electronica. 14(3): 19A, 12p.  palaeo-electronica.org/2011_3/1_adams/index.html

79    Fiorillo, A.R.  2011.  Microwear patterns on the teeth of northern high latitude hadrosaurs with comments on microwear patterns in hadrosaurs as a function of latitude and seasonal ecological constraints. Palaeontologia Electronica. 14(3): 20A, 17p.  palaeo-electronica.org/2011_3/7_fiorillo/index.html

80    Chinsamy, A., Thomas, D.B., Tumarkin-Deratzian, A.R., and Fiorillo, A.R. 2012. Hadrosaurs were perennial polar residents.  The Anatomical Record 295:610-614.

Manuscripts in Press

81    Fiorillo, A.R., and Tykoski, R.S.  in press.  A new species of centrosaurine ceratopsid Pachyrhinosaurus from the North Slope (Prince Creek Formation: Maastrichtian) of Alaska.  Acta Palaeontologica Polonica.

82    Fiorillo, A.R., Adams, T.L., and Kobayashi, Y., 2012, New sedimentological, palaeobotanical, and dinosaur ichnological data on the palaeoecology of an unnamed Late Cretaceous rock unit in Wrangell-St. Elias National Park and Preserve, Alaska, USA: Cretaceous Research, doi: 10.1016/j.cretres.2012.04.013.

83    Flaig, P.P., McCarthy, P.J., and Fiorillo, A.R.  in press.  Anatomy, evolution and paleoenvironmental interpretation of an ancient Arctic coastal plain: Integrated paleopedology and palynology from the Upper Cretaceous (Maastrichtian) Prince Creek Formation, North Slope, Alaska, USA.  In, New Frontiers in Paleopedology and Terrestrial Paleoclimatology. Driese, S. and Nordt, L. (eds.), SEPM Special Publication.

84    Fiorillo, A.R., and Adams, T.L. in press.  A therizinosaur track from the Lower Cantwell Formation (Upper Cretaceous) of Denali National Park, Alaska.  PALAIOS.

Popular and Non-technical Publications

85       Fiorillo, A.R., Harris, R., and May, C.L.  1996.  Late Jurassic (Morrison Formation) dinosaur remains from Curecanti National Recreation Area, Colorado.  Park Science. 16(4):14-15.

86       Fiorillo, A.R., May, C.L., and Harris, R.  1997.  Comparison of Paleontological Resource Management in the Rocky Mountain Region by the National Park Service and the U.S. Forest Service.  In, Making Protection Work, Proc. Ninth Conf. Research and Resource Management in Parks and on Public Lands.  Harmon, D. (ed.), The George Wright Society, Hancock, MI, Pp. 308-311.

87    Fiorillo, A.R., and Harris, R.L. 2000.  Late Jurassic (Morrison Formation) continental trace fossils from Curecanti National Recreation Area, Colorado.  Park Science. 20(2):28-29, 35.

88    Fiorillo, A.R., and Santucci, V.L. 2001.  Crossing park boundaries in the study of ancient ecosystems.  In, Crossing Boundaries in Park Management, Proc. 11th Conf. Research and Resource Management in Parks and on Public Lands.  Harmon, D. (ed.), The George Wright Society, Hancock, MI.  Pp. 365-370.

89    Fiorillo, A.R.  2002.  Preliminary report on recent discoveries during a paleontological survey in Aniakchak National Monument and Preserve – 2001.  Park Paleontology.  National Park Service Geologic Resources Division, Paleontology Program. 6(1):5.

90    Fiorillo, A.R.  2002.  Remains of the last of the sauropod dinosaurs excavated at Big Bend National Park.  Park Paleontology.  National Park Service Geologic Resources Division, Paleontology Program. 6(1):6.

91    Fiorillo, A.R.  2002. Taphonomy: the story of death and destruction.  Pp. 109-116. In, Dinosaurs: the science behind the stories.  Scotchmoor, J.G., Springer, D.A., Breithaupt, B.H., and Fiorillo, A.R. (eds.).  American Geological Institute, Alexandria.

92    Fiorillo, A.R., Armato, P., and Kucinski, R.  2004.  Wandering rocks in Kenai Fjords National Park.  Alaska Park Science, 3:21-23.

93    Fiorillo, A.R., Kucinski, R., and Hamon, T.  2004.  New Frontiers, Old Fossils: recent dinosaur discoveries in Alaska National Parks.  Alaska Park Science, 3:4-9.
94    Fiorillo, A.R., 2004.  The dinosaurs of arctic Alaska.  Scientific American. v. 291, no. 6, p. 84-91.

95    Fiorillo, A.R., 2007.  A dinosaur discovery in Aniakchak National Monument.  In, Beyond the Moon Crater Myth: a new history of the Aniakchak Landscape, Ringsmuth, K.J. (ed.), National Park Service, Research/Resources Management Report AR/CRR-2207-63,  p. 230-231.

Dr. Tykoski's recent papers

  1. The Pachyrhinosaurus perotorum manuscript of Dr. Fiorillo and Dr. Tykoski can be found here.
  2. Large Dinosaur Excavated for Scientific Study
  3. Dinosaur Excavation: Latest Photos of the Big Bend Giant
  4. Tykoski, R. S., T. B. Rowe, R. A. Ketcham, and M. W. Colbert. 2002. Calsoyasuchus valliceps, a new crocodyliform from the Early Jurassic Kayenta Formation of Arizona. Journal of Vertebrate Paleontology 22:593-611.
  5. Tykoski, R. S., C. A. Forster, T. Rowe, S. D. Sampson, and D. Munyikwa. 2002. A furcula in the coelophysid theropod Syntarsus. Journal of Vertebrate Paleontology 22:728-733.
  6. Tykoski, R. S., and T. Rowe. 2004. Ceratosauria; pp. 47-70 in D. B. Weishampel, P. Dodson, and H. Osmolska (eds.), The Dinosauria, Second Edition. University of California Press, Berkeley, California.
  7. Tykoski, R. S. 2005. Vertebrate paleontology in the Arizona Jurassic; pp. 72-93 in R. D. McCord (ed.), Vertebrate Paleontology of Arizona. Mesa Southwest Museum Bulletin Number 11. Mesa, Arizona.
  8. Tykoski, R. 2007. Small theropods, big ideas; pp. 86 in T. Holtz, Jr., Dinosaurs: the most complete, up-to-date encyclopedia for dinosaur lovers of all ages. Random House, New York, New York.
  9. Tykoski, R. S., and A. R. Fiorillo. 2010. An enantiornithine bird from the lower Middle Cenomanian of Texas. Journal of Vertebrate Paleontology 30:288-292.
  10. Rowe, T., R. Tykoski, and J. Hutchinson. 1997. Ceratosauria; pp. 106-110 in P. J. Currie and K. Padian (eds.), Encyclopedia of Dinosaurs. Academic Press, San Diego, California.
  11. Allain, R., R. Tykoski, N. Aquesbi, N.-E. Jalil, M. Monbaron, D. Russell, and P. Taquet. 2007. An abelisauroid (Dinosauria: Theropoda) from the Early Jurassic of the High Atlas Mountains, Morocco, and the radiation of ceratosaurs. Journal of Vertebrate Paleontology 27:610-624.
  12. Fiorillo, A. R., R. S. Tykoski, P. J. Currie, P. J. McCarthy, and P. Flaig. 2009. Descritpion of two partial Troodon braincases from the Prince Creek Formation (Upper Cretaceous), North Slope Alaska. Journal of Vertebrate Paleontology 29: 178-187.
  13. Fiorillo, A. R., and R. S. Tykoski. In press. A new species of the centrosaurine ceratopsid Pachyrhinosaurus from the North Slope (Prince Creek Formation: Maastrichtian) of Alaska. Acta Palaeontologica Polonica, doi: here.